Read e-book online Adaptation in Stochastic Environments PDF
By Jin Yoshimura, Colin W. Clark
The classical concept of average choice, as constructed via Fisher, Haldane, and 'Wright, and their fans, is in a feeling a statistical concept. normally the classical concept assumes that the underlying surroundings during which evolution transpires is either consistent and good - the idea is during this feel deterministic. in truth, nevertheless, nature is nearly continually altering and risky. we don't but own a whole idea of ordinary choice in stochastic environ ments. maybe it's been inspiration that this kind of idea is unimportant, or that it might be too tough. Our personal view is that the time is now ripe for the advance of a probabilistic conception of usual choice. the current quantity is an try to offer an common advent to this probabilistic concept. each one writer used to be requested to con tribute an easy, easy creation to his or her strong point, together with energetic discussions and hypothesis. we are hoping that the ebook contributes extra to the knowledge of the jobs of "Chance and Necessity" (Monod 1971) as built-in elements of variation in nature.
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Extra info for Adaptation in Stochastic Environments
Strategy 1 emerging here in low frequency will be excluded if (21) which is satisfied whenever (22) Since the second term on the right hand side is positive for YeN) > 0, conditions for exclusion of the rare strategy (23) always can be satisfied by choosing Kl not too much higher than K 2 . This model version is illustrated on Fig. 6b. The only difference in this figure compared to Fig. 6a is that environmental fluctuation acts on the position of the established population of strategy 2. If the variance in density is high enough, both of the strategies become unable to invade the other one: the rare strategy is excluded.
Let us choose the parameters of strategy 1 (a, b, K 1 ) according to (10) and an arbitrary distribution for the environmental parameter ~(t). These determine Nl and V(N) for the population of strategy 1. Clearly the result is independent of parameters of strategy 2 and satisfies condition (16). Next we can choose K2 within the interval between Nl and K 1 , which guarantees that both of the conditions in (18) are satisfied. c may be any positive number. With these parameters the two strategies are coexisting, while if K2 is chosen out of the interval [N 1, K 1 ], coexistence becomes impossible and the strategy with the higher equilibrium density outcompetes the other one.
Bioi. 12:119-129. Cohen, D. 1967. Optimising reproduction in a randomly varying environment when a correlation may exist between the conditions at the time a choice has to be made and the subsequent outcome, J. Theoret. Bioi. 16:1-14. Cohen, D. and S. Levin. 1987. The intreraction between dispersal and dormancy strategies in varying and heterogeneous environments. In Lecture Notes in Biomathematics (eds. Teramoto, E. ) 71:110-122. P. 1985. ESS germination strategies in randomly varying environments.
Adaptation in Stochastic Environments by Jin Yoshimura, Colin W. Clark